Type Material
Holotype: “Male holotype and 2 male paratypes from Twenty-nine Palms, California, USA (AMNH). Female allotype from 12 km northwest of Las Vegas, Nevada, USA (AMNH). We were able to examine the male holotype and female allotype” (Cushing et al., 2018, p. 460).
Measurements: “Males (n = 16). TL 24.0–33.5; CL 7.1–10.4; CH 3.0–6.2; FNL 0.3–0.5; FNH 0.4–0.7; FFH 0.6–0.9; PL 20.0–35.0; PT 1.0–4.1; PMT 5.1–10.0; LI 12.0–25.0; LIV 24.0–33.0; PPL 4.0–5.4; A/CP 4.1–7.8; FNL/FNH 0.5–1.0; FFH/FNH 1.0–2.0; FFH/CH 0.1–0.3” (Cushing et al., 2018, p. 461).
“Females (n = 17). TL 27.0–48.0; CL 8.6–13.9; CH 4.0–5.9; PL 20.0–37.0; PT 1.7–2.0; PMT 4.6–6.9; LI 12.0–24.5; LIV 25.5–41.0; PPL 3.9–6.4; A/CP 4.5–6.3” (Cushing et al., 2018, p. 461).
Operculum Description: “Muma’s illustration is accurate (1951, fig. 43). Club-shaped with long arms ending in oval shaped wings (Fig. 4F)” (Cushing et al., 2018, p. 461).
Chelicerae Description: “Male. Deep VDC, similar to E. calexicensis but with slightly mesal orientation (Fig. 1G). Fixed finger slightly thicker than E. calexicensis, with no median dentition. Movable finger: MP-2 MSM-MM; distinct MPL (Figs. 2W & X). Fondal notch tiny with one RFA and two-three RFA on the ventral region of fixed finger; fondal teeth graded I, III, II, IV retrolaterally; I, II, III, IV prolaterally (Fig. 3G)” (Cushing, et al., 2018, p. 461).
“Female. Fixed finger: FP-1 FSM-FM-FSD-FD. Movable finger: MP-2 MSM, proximal larger-MM; distinct MPL (Figs. 2Y & Z). One large RFA; fondal teeth graded II, I, III, IV retrolaterally, and I, II, tiny III, serrate IV prolaterally” (Cushing et al., 2018), p. 461).
Diagnosis: “Eremocosta titania is related to E. calexicensis but differs in coloration, especially of appendages; fixed finger longer and narrower than E. calexicensis, MP of movable finger more robust as well (compare Figs. 2W & X with Figs. 2E & F). Male E. titania are also generally smaller than male E. calexicensis. Female genital operculum club-shaped with long arms; distinct from operculum of E. titania. See also diagnosis above under E. calexicensis” (Cushing et al., 2018, p. 461).
Other Information: “Specimens from the more northern regions, Nevada, Northern California have a darker coloration than those found in regions of Southern California but male cheliceral dentition and female genital operculum remains relatively constant” (Cushing et al., 2018, p. 462).
Eremocosta titania – male holotype
AMNH: Twentynine Palms, California; July – Aug., 1945; coll. Jefferson H. Branch
Originally IDd as Eremorhax titania Muma by Martin H. Muma, 1950
Measurements:
| Char | Character description | M (mm) |
| CL | chelicerae length | 9.20 |
| CH | Chelcera height | 4.25 |
| FNL | Fondal notch length | 0.22 |
| FNH | Fondal notch height | 0.73 |
| FFH | Fixed finger height | 0.81 |
| LI | First leg length | 27.00 |
| LIV | Fourth leg length | 42.20 |
| PPL | Propeltidium length | 4.45 |
| PPW | Propeltidium width (from bottom edge where “ears” begin) | 6.40 |
| PL | Palpus length | 35.50 |
| PMT | Palpal metatarsus length | 8.30 |
| PT | Palpal tarsal length | 1.67 |
| TL | Total length | 41.00 |
Notes:
Took photo of fondal teeth of left chelicerae since I could more easily open the fingers of this chelicerae.
Took later photo of fondal teeth right chelicerae since I figured out how to do this!
Pedipalp – no color, no papillae
