Tyloderma sphaerocarpae Wibmer, 1981
Source: O'Brien, C.W., Wibmer, G.J. 1982.
Family: Curculionidae
Tyloderma sphaerocarpae image
Michael Jansen  
12. Tyloderma sphaerocarpae Wibmer, new species Holotype female.-Body elongate; cuticle reddish brown, elytra with darker spots and bands; with short, slender, yellowish white, recumbent scales generally forming spots or bands. Rostrum with more or less triangular, median, impunctate area, between 1/2 and 3/4 from base; punctures medium sized, moderately deep, very dense on basal l/2, on apical 1/2 small to medium sized, shallow to moderately deep, moderately dense to moderately sparse, larger and deeper toward sides. Head with few scales above eyes and on middle of vertex; with short, deep, longitudinal frontal sulcus; punctures medium sized, moderately deep, dense above eyes, sparser toward middle, much shallower toward vertex; eyes moderately convex. Prothorax l/20 longer than wide, moderately constricted on flanks near apex, sides diverging moderately in basal 1/3, converging strongly to constriction, then moderately toward apex; pronotal disc on each side with lateral band of scales on basal 2/5, subparallel to side, and spot ca. 3/5 from base, closer to midline, few scales on midline on apical 1/4, and few subapical along margin; punctures medium to large, deep and dense, generally separated by 1/2 their own diameters; larger, deeper and denser along sides of pronotum and on flanks. Elytra 1.57 times as long as wide, 2.02 times as long as prothorax, sides diverging very weakly in basal 2/5, converging weakly to declivity, then more strongly toward apex; scales generally moderately sparse; strial punctures on basal 1/3 of disc large, deep, dense, generally separated by their own diameters, on striae 9 & 10 irregularly shaped, shallower, more uneven; strial grooves 1 to 8 distinct on apical 1/2, 9 very distinct on apical 2/3; intervals moderately sinuate at base, moderately convex to convex on basal 2/3, strongly convex on apical 1/3, 4 to 8 moderately raised at their apices. Mesosternum with lateral processes ca. 3/5 from anterior margin, moderately distinct, posterior margin moderately narrow; surface moderately convex along midline, with some scales on posterior 1/2, punctures medium sized, moderately shallow, moderately dense, each with short, fine, subrecumbent to suberect seta; mesepisternum with few, medium sized, deep, mostly contiguous punctures forming longitudinal sulcus. Metasternum more or less flat on median area; punctures medium to large, moderately deep, moderately dense on median area, on lateral areas larger, deeper and denser. Abdomen with sterna 1 &: 2 moderately convex, sterna 3 &: 4 longitudinally nearly flat; punctures small, moderately shallow, moderately sparse (larger, deeper and denser between metacoxae and along base) on sternum 1, on sternum 2 small to medium sized, shallow to moderately shallow, moderately sparse to sparse, on sterna 3 &: 4 small, moderately deep, dense, most in transverse row, on sternum 5 medium sized, moderately deep, moderately dense to dense. Legs primarily reddish brown, with trochanter, base and apex of femur, apex of tibia, and tarsus darker; femur with inner margin weakly rounded, surface strongly rugosely striate; tibia moderately rugosely striate; praemucro moderately oblique to axis of tibia and directed away from uncus. Genitalia: fig. 109. Length (pronotum + elytra): 4.45 mm. (1.475 + 2.975). Allotype male.-Metasternum weakly concave on median area; abdominal sternum 1 concave along midline, sternum 2 very weakly convex; uncus moderately oblique to axis of tibia, arising from middle of apex, praemucro ca. 1/7 from apex of tibia on fore- and midlegs, ~• 1/5 from apex on hind legs (as in fig. 59). Genitalia: fig. 77. Length (pronotum + elytra): 4.05 mm. (1.25 + 2.80). Intraspecific variatim.-The venter is often darker than the dorsum, at least in part; and the scales are yellowish white to yellow. The rostrum is 1.81-2.25 times as long as its width at the apex, with its impunctate median area sometimes weakly distinct. The punctures on the apical 1/2 are moderately dense to dense. There are a few scales at the base of the rostrum and around the eyes, and sometimes these are more or less contiguous forming a transverse dorsal band on the vertex. The frontal sulcus varies from scarcely distinct, to distinct and long, reaching the vertex. The punctures above the eyes sometimes are moderately shallow, not sparser toward the middle of the head. The prothorax has a ratio (length/width) of 1.00-1.11 (1.06 ± 0.03), its widest point located between 1/4 and 2/5 from the base. The elytra are 1.54-1.76 (1.64 ± 0.05) times as long as wide, and 1.94-2.22 (2.08 ± 0.08) times as long as the prothorax, with their widest point located between 1/4 from the base and near the middle. Their sides sometimes converge very weakly from the humeri to the declivity. The strial punctures are denser on some specimens, generally separated by 1/2 to 3/4 their diameters. Sometimes they are moderately distinct on the apical 1/3 of the 9th stria; and the intervals 4 to 8 may be only moderately raised at their apices. The punctures on abdominal sternum 1 are sometimes medium sized, moderately deep, and moderately dense. Rarely, they do not form a clear transverse row on sternum 4. The inner margin of the femur is sometimes moderately rounded, and in some specimens it is obtusely angulate. Length, pronotum + elytra: 3.15-4.80 mm. (4.01 ± 0.33). Remarks and comparative notes.-In the Variegatum group this species is distinguished from all but aquaticum by the more elongate form and comparatively smaller prothorax. It can be separated from aquaticum by its usually larger size, more convex intervals, darker abdominal sterna with larger and deeper punctures, and by the usually denser scale pattern of the elytra. It differs from variegatum by the usually straighter intervals and lighter color. It has the elytral strial punctures on the basal 1/4 much larger than the pronotal punctures, and more convex intervals, which readily separate it from rufescens. Etymology.-species named for its probable host plant species, Ludwigia sphaerocarpa. Type Locality.-U.S.A., Florida, Wakulla County, 1 mile north of Spring Creek.